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Gopher tortoise at St

Animal

Gopher tortoise

The gopher tortoise is a land turtle with shovel-like front legs. It digs long tunnels in sandy ground. Many other animals use its tunnels too!

The gopher tortoise lives in the southeastern United States. With its strong, shovel-like front legs it digs deep burrows in dry, sandy soil, and more than 350 other kinds of animals share them. It eats low-growing plants. Losing its habitat has made it critically endangered.

Gopherus polyphemus is a burrowing tortoise (family Testudinidae) of the southeastern United States, from southern South Carolina through most of Florida west to eastern Louisiana. A keystone species and ecosystem engineer, its burrows may house over 350 other species. Stemle and Howell assessed it as Critically Endangered (A2bcde+3bce+4bce) for the IUCN Red List (assessed 7 August 2024, published 2026), with an estimated reduction of at least 80% over the past three generations (120 years) and a projected further decline of more than 95%.

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Photo: © Hans Hillewaert, CC BY-SA 4.0, via Wikimedia Commons

Meet the gopher tortoise

The gopher tortoise is a turtle that lives on land.

It has a brown, domed shell.

It is a great digger!

The gopher tortoise is a land turtle from the southeastern United States. Its shell is oblong and usually brown, tan or gray, and an adult's shell is on average about 23 to 28 centimeters long. A typical adult weighs about 5.5 kilograms.

It is a strong digger. Its front legs are flat and shovel-like, and its back feet are flat and stumpy, with no webbing between the toes.

Gopherus polyphemus belongs to the tortoise family, Testudinidae. The Animal Diversity Web (University of Michigan) gives an average adult carapace length of 23 to 28 centimeters, a maximum of about 38.7 centimeters and an average mass of 5.5 kilograms (from the AnAge database), and describes an oblong brown, tan or gray carapace, an unhinged dull yellow and pale brown plastron, flattened, stump-like, unwebbed hind feet and shovel-like forelimbs adapted for digging.

The IUCN notes that there are no described subspecies, and that the US Fish and Wildlife Service treats the western population (Louisiana, Mississippi and western Alabama) as a distinct population segment for listing purposes.

Gopher tortoise at St
Photo: © Hans Hillewaert, CC BY-SA 4.0, via Wikimedia Commons

Quick facts

Group
Reptile
Length
0.23 to 0.28 m
Weight
5.5 kg

Sandy pinelands of the Southeast

Gopher tortoises live where the ground is dry and sandy.

They like open pine woods with lots of low plants.

Gopher tortoises live in the southeastern United States, from southern South Carolina through most of Florida and west to eastern Louisiana. They need dry, sandy soil that is easy to dig, open woods with a thin tree canopy, and plenty of low plants to eat.

Their homes include longleaf pine savannas, sandhills, scrub, flatwoods and coastal dunes. They also turn up in old pastures, fields and grassy roadsides, and in Florida even in people's yards.

The IUCN describes a range from southern South Carolina through most of Florida and west along the coastal plain to eastern Louisiana, from near sea level to about 200 meters, now highly fragmented by development. Natural habitats include sandhill, scrub, flatwoods, dry prairie, xeric hammock, coastal strand and longleaf pine savanna, and tortoises often use grassy open areas, old pastures, fields, roadsides and utility rights-of-way; across all of them they need well-drained soils for burrows and enough groundcover to feed on. The Florida Fish and Wildlife Conservation Commission lists forests, pastures and yards.

The Animal Diversity Web adds that periodic burning is an essential part of the habitat, because it holds back the tall, dense vegetation that would shade out the low-growing plants tortoises eat.

Continent

Where can you find this animal?

  • Lives here naturally · Countries and seas named by a scientific source
  • Sightings recorded · Records of wild sightings (GBIF)
Lives here naturally · Countries and seas named by a scientific source

The IUCN lists gopher tortoises only in the southeastern United States: Alabama, Florida, Georgia, Louisiana, Mississippi and South Carolina, from southern South Carolina through most of Florida and west to eastern Louisiana. The whole country is shaded, but they live only in this corner of it.

Places: United States.

Source: iucn-red-list · checked October 2026

Sightings recorded · Records of wild sightings (GBIF)

Counts of GBIF records of wild observations (human and machine) and preserved specimens. Records include introduced populations, vagrants, escaped or released animals and misidentifications, so a country with records is not necessarily part of the native range.

Wild sightings have been recorded in United States, Belize, Denmark, Mexico and Colombia.

Source: gbif · checked October 2026

Range maps show where scientists say an animal lives, often country by country. An animal may live in only part of a country shown, and sightings records can include animals that wandered far from home.

Grazing on low plants

Gopher tortoises eat plants.

They munch grasses, leaves and fruit.

Gopher tortoises are plant eaters. They graze on low-growing plants: grasses, small flowering plants, leaves, fruits and even cactus. Scientists have recorded them eating 69 different kinds of plants.

By eating fruits and other plants, they help spread seeds, which can increase the variety of plants growing in an area.

The IUCN describes the gopher tortoise as usually a vegetarian dietary generalist, eating grasses, forbs, fruits, cactus and leaves, with 69 plant genera and species documented (Macdonald and Mushinsky 1988), varying with habitat. Tortoises are well documented as consumers and seed dispersers of native and non-native plants, which can increase plant diversity.

The Florida Fish and Wildlife Conservation Commission summarizes the diet as low-growing plants, and the Animal Diversity Web links healthy feeding habitat to regular fire, which keeps the groundcover open.

Did you know?

More than 350 kinds of animals use gopher tortoise burrows.

Gopher tortoise burrows are shared by more than 350 other kinds of animals.

Burrows may house over 350 invertebrate and vertebrate commensal species (IUCN, after Eisenberg 1983; FWC).

Built for digging

Its front legs are flat like shovels.

Its back feet are flat and stumpy.

Its shell is hard and domed.

A gopher tortoise is built to dig. Its front legs are flattened like shovels, and its back feet are flat and stumpy.

Young tortoises have growth rings on their shell scutes, which can help estimate their age, but on old tortoises the shell wears smooth. Males and females look alike, but a male's bottom shell curves inward a little, while a female's is flatter.

The Animal Diversity Web describes shovel-like forelimbs adapted for digging and flattened, stump-like, unwebbed hind feet; an oblong carapace whose scutes show growth rings in younger tortoises (usable to estimate age) but wear smooth in older ones; and hatchlings with yellow-centered, brown-bordered scutes and a yellow or orange plastron.

Adult males and females are similar in size and appearance, though females may average slightly larger. The most reliable difference is the male's concave plastron (the female's is flatter); males also have larger mental glands on the chin and longer gular projections at the front of the plastron (Mushinsky et al. 1994, via ADW).

Tap a part to find out more

How big is it?

Length: 0.23 to 0.28 m (average adult shell (carapace) length about 23 to 28 centimeters)

Weight: 5.5 kg (average adult weight about 5.5 kilograms)

Eggs in the sand, slow to grow up

The mother lays about six eggs in the sand.

The babies are very small.

Gopher tortoises can live a very long time.

A female lays no more than one clutch of eggs a year, usually about six eggs, in sandy soil. In Florida the eggs hatch after about 67 days; in Mississippi, it takes about 88 days.

The hatchlings are only about 5 centimeters long, and many eggs and babies are eaten by predators. Survivors grow slowly: they are not ready to breed until they are about 9 to 20 years old. Gopher tortoises can live for more than 70 years.

The Animal Diversity Web gives a single clutch a season, averaging six eggs (range five to nine), temperature-dependent sex determination with an unusually low pivotal temperature of about 29 °C (Burke et al. 1996), and very high nest and hatchling predation. The IUCN summarizes clutch sizes averaging 6.4 across studies (study means 3.8 to 11.1), incubation of about 88 days in Mississippi (range 52 to 109) and 67 days in Florida (the Animal Diversity Web gives 80 to 100 days, shorter farther south), a hatchling carapace length of about 48 millimeters in Mississippi, sexual maturity at about 9 to 20 years depending on latitude and habitat, a generation length of about 40 years or more, and longevity that can exceed 70 years.

One study (Alford 1980) recorded more than 94% mortality of eggs and hatchlings, and another (Landers et al. 1982) estimated that a female produces a successful clutch only about once every 9 to 10 years (IUCN).

A life, step by step

A burrow full of neighbors

Gopher tortoises dig long tunnels called burrows.

More than 350 kinds of animals use these burrows too.

The burrows keep animals safe from fire and heat.

Gopher tortoises dig burrows that are on average about 4.5 meters long and 2 meters deep. Inside, the temperature and dampness stay steady all year, so the burrow protects the tortoise from fire, heat, cold and drought.

The Florida Fish and Wildlife Conservation Commission says more than 350 other kinds of animals share gopher tortoise burrows. That is why the tortoise is called a keystone species: many other living things depend on what it does.

The Animal Diversity Web gives an average burrow length of 4.5 meters and depth of 2 meters, with a steady temperature and humidity through the year that protects the tortoise from fires, extreme temperatures and drought. The IUCN calls the species a keystone species and ecosystem engineer: its expansive burrows may house over 350 invertebrate and vertebrate commensals (Eisenberg 1983), its digging aerates the soil, and its burrows give other animals refuge during wildfires, prescribed fires and extreme temperatures.

ADW notes that individuals defend overlapping territories, with ramming and charging seen more often in the breeding season, and the IUCN notes that where habitat is not burned, tortoises tend to move to road and utility rights-of-way for food and basking, which raises their risk from traffic.

Did you know?

Scientists have seen them eat 69 kinds of plants.

Gopher tortoises have been recorded eating 69 different kinds of plants.

69 plant genera and species documented in the diet (Macdonald and Mushinsky 1988, via IUCN).

Critically endangered

Gopher tortoises are in great danger.

Many of their sandy homes have been built on.

People protect them and move them to safe places.

The gopher tortoise is critically endangered. Most of the longleaf pine forest it lives in is gone: the IUCN says it has shrunk by 95 to 97% from its original size. Towns, roads, farms and mines also destroy and split up its habitat, and without regular fires many open woods grow too thick.

Predators such as raccoons, and invaders such as fire ants and feral hogs, eat many eggs and babies. In Florida the tortoise and its burrow are protected by law, and tortoises must be moved to safety before land is cleared.

Stemle and Howell assessed Gopherus polyphemus as Critically Endangered (A2bcde+3bce+4bce) for the IUCN Red List (assessed 7 August 2024, published 2026): an estimated reduction of at least 80% over the past three generations (120 years), a projected reduction of more than 95% over the next three, and an ongoing 80% reduction across two past generations and one future generation. In Florida, populations fell 50 to 80% since the 1960s. Threats include habitat loss and fragmentation from urbanization, roads, mining, agriculture and solar development; too little or badly timed fire; invasive predators (red imported fire ants, tegus, feral hogs) and subsidized predators; disease; and past hunting for food in the 20th century. Total protected population size was estimated at 70,122 in 2022 (Folt et al. 2022). It was previously assessed as Vulnerable (1996).

The western population has been federally listed as Threatened under the US Endangered Species Act since 1987 (IUCN). The Florida Fish and Wildlife Conservation Commission lists the species as Threatened, protects both tortoise and burrow, and requires permits and relocation before land clearing.

IUCN Red List

How is the gopher tortoise doing in the wild?

  1. Least Concern
  2. Near Threatened
  3. Vulnerable
  4. Endangered
  5. Critically Endangered (this animal)
  6. Extinct in the Wild
  7. Extinct

Critically Endangered

At extremely high risk of disappearing from the wild.

See the IUCN Red List assessment

Amazing facts

  • More than 350 kinds of animals use gopher tortoise burrows.

    Gopher tortoise burrows are shared by more than 350 other kinds of animals.

    Burrows may house over 350 invertebrate and vertebrate commensal species (IUCN, after Eisenberg 1983; FWC).

  • A gopher tortoise burrow is about as long as a car.

    An average burrow is about 4.5 meters long and 2 meters deep.

    Average burrow length 4.5 m and depth 2 m (ADW).

  • Gopher tortoises can live more than 70 years.

    A gopher tortoise can live for more than 70 years.

    Longevity can exceed 70 years (IUCN).

  • Scientists have seen them eat 69 kinds of plants.

    Gopher tortoises have been recorded eating 69 different kinds of plants.

    69 plant genera and species documented in the diet (Macdonald and Mushinsky 1988, via IUCN).

Words to know

burrow
a tunnel or hole an animal digs to live in
keystone species
an animal or plant that many other living things depend on
carapace
the top part of a turtle's shell
clutch
a group of eggs laid at one time
longleaf pine
a tall pine tree of the southeastern United States that grows in open, often-burned woods
critically endangered
at extremely high risk of dying out in the wild

Discovery challenge

  1. What does a gopher tortoise dig?

    What does a gopher tortoise dig with its front legs?

    What structure makes the gopher tortoise an ecosystem engineer?

    • A long burrow
    • A nest in a tree
    • A pond
    Show the answer

    A long burrow

    A burrow!

    A burrow about 4.5 meters long and 2 meters deep, on average.

    Its expansive burrows, used by over 350 other species (IUCN; ADW).

  2. How many kinds of animals share its burrows?

    About how many other kinds of animals use gopher tortoise burrows?

    How many commensal species may use gopher tortoise burrows?

    • More than 350
    • About 5
    • None
    Show the answer

    More than 350

    More than 350!

    That is why it is called a keystone species.

    Over 350 invertebrate and vertebrate commensals (IUCN; FWC).

  3. What does a gopher tortoise eat?

    What kind of food does a gopher tortoise eat?

    How does the IUCN describe the gopher tortoise's diet?

    • Plants
    • Fish
    • Insects only
    Show the answer

    Plants

    Plants!

    Grasses, leaves, fruits and even cactus.

    Usually a vegetarian dietary generalist (IUCN).

  4. What kind of ground does a gopher tortoise need?

    What kind of soil do gopher tortoises need for their burrows?

    Which soil property limits where gopher tortoises can live?

    • Dry, sandy soil
    • Frozen ground
    • Wet mud under water
    Show the answer

    Dry, sandy soil

    Dry, sandy ground!

    Dry, sandy soil that is easy to dig.

    Well-drained soils for burrow construction (IUCN; ADW).

Places where you might see it

Explore next

For parents & teachers

What children will learn

  • Describe how a gopher tortoise's body suits digging.
  • Explain what a keystone species is, using the gopher tortoise's burrows.
  • Explain why fire matters for gopher tortoise habitat.
  • Name the main threats to gopher tortoises and ways people protect them.

Recommended ages: 3 to 17 · About 15 minutes

Talk about it

  • Why might so many animals use a tortoise's burrow?
  • How could a forest growing too thick be bad for a tortoise?
  • Why might a tortoise that grows up slowly be harder to protect?

Try at home or in class

  • Burrow measure: with a grown-up, measure 4.5 meters on the floor or sidewalk with a tape measure to see how long an average gopher tortoise burrow is.
  • Shovel test: try scooping dry sand with a flat spoon and with a pointed stick, and compare which moves more sand, like a tortoise's flat front legs.

Key vocabulary: burrow, keystone species, carapace, clutch, longleaf pine, critically endangered

Where these facts come from

  1. GBIF Backbone Taxonomy and GBIF Occurrence Search , Global Biodiversity Information Facility (CC0 1.0 (backbone); occurrence counts are facts derived from records under CC0, CC BY and CC BY-NC)
  2. Wikidata , Wikimedia Foundation (CC0 1.0)
  3. The IUCN Red List of Threatened Species , International Union for Conservation of Nature (IUCN terms of use (no redistribution of Red List data or range maps))
  4. Gopherus polyphemus (Stemle and Howell, assessed 7 August 2024, published 2026) , IUCN Red List of Threatened Species
  5. Gopherus polyphemus: justification, range, population, habitat, threats and conservation text , IUCN Red List of Threatened Species
  6. Gopherus polyphemus: IUCN Red List global assessment record (Stemle and Howell 2026), IUCN checklist on GBIF , GBIF
  7. Gopherus polyphemus (gopher tortoise) , Animal Diversity Web, University of Michigan Museum of Zoology
  8. Gopher tortoise , Florida Fish and Wildlife Conservation Commission
  9. File:Gopherus polyphemus (side).jpg (8 May 2011, Hans Hillewaert) , Wikimedia Commons
  10. Wikimedia Commons , Wikimedia Foundation (Per file: Public domain, CC0, CC BY or CC BY-SA only (credited on each page))

Checked, not yet human-reviewed. Last updated November 1, 2026. Spotted a mistake? Tell us.

Picture credits
  • Photo: © Hans Hillewaert, CC BY-SA 4.0, via Wikimedia Commons · source · license